
Sunday, 30 September 2007
Quiz Woodpecker 5

Friday, 28 September 2007
Woodpeckers: juvenile moult
Juvenile woodpeckers begin a partial post juvenile moult at around the time of fledging and this continues until the autumn. In this first moult, juveniles of all ten European woodpecker species replace their primaries, rectrices and overall (body) plumage but some tertials, secondaries, primary coverts and outer greater coverts are retained. One of the ways to age 1st winter birds is the fact that there is often a moult limit in the greater coverts. For example, in Great Spotted Woodpecker where the new adult-type black inner greater coverts, contrast with the old juvenile brownish outer greater coverts. Juveniles have shorter tails than adults, due to all but the central pair of rectrices being shorter than on adults. The odd looking stunted tail of juvenile Black Woodpeckers in flight is due to this tail arrangement. Another interesting case involves Three-toed Woodpecker. Juveniles of this species sometimes begin their moult while still in the nesting cavity. Interestingly, the fifth to seventh innermost primaries are grown and replaced before the birds have fledged and thus they have never been used. This apparent rush to moult is almost certainly an adaptive feature, the result of the need of the species, as a bird of mainly northern latitudes, to complete the moulting task before the end of the short summer season. When the harsh winter season begins birds must be ready to disperse at short notice to new foraging areas, often some distance away, and a delay while moult is completed could prove fatal. For similar reasons, suspended moult occurs in local populations of other species that are prone to eruptive movements such as Fenno-Scandic and Russian Great Spotted Woodpeckers. Juveniles trapped in Fenno-Scandia after arriving from further east in late autumn apparently often exhibit signs of suspended moult in the tail and wing when compared to resident birds. Moult is then resumed sometimes as late as December.
Woodpecker moult
The moult regime of the true woodpeckers is rather unusual and this is related to the lifestyle of the family, in particular to foraging habits and to the breeding cycle. A typical European woodpecker (all except Wryneck) has ten primaries, eleven secondaries, and twelve tail feathers. Primaries 6, 7 and 8 are the longest and P10 is reduced in size in adults but is longer and broader in juveniles. Most adult woodpeckers begin a complete post-breeding moult in late spring or early summer soon after breeding has finished and this lasts into autumn. Primaries are moulted sequentially (ascendant) from P1 and the secondaries from two centres (ascendantly from S1, ascendantly and descendantly from S8). The outermost rectrices are also much shorter than the other tail feathers, the central two being the longest. These two feathers are so important as props when climbing and clinging to vertical surfaces, so they are replaced last, after the other tail feathers. Counting the central two as R1 moult starts ascendantly in pairs from R2 to R6 with R1 last. There is no pre-breeding moult for the true woodpeckers, though Wryneck has a partial pre-breeding moult. Wrynecks moult their tail feathers in reversed sequence.
Wednesday, 26 September 2007
Monday, 24 September 2007
Holes: White-backed Woodpecker - natural hole use

Holes: White-backed Woodpecker

Holes: secondary user - Tengmalm's Owl

Holes: Black Woodpecker as a provider

Saturday, 22 September 2007
Holes: location

Holes: used and unused

Holes: safety

.
Friday, 21 September 2007
Holes

Green Woodpecker: photo of male
Thursday, 20 September 2007
Quiz Woodpecker 2
Wednesday, 19 September 2007
Lesser Spotted Woodpecker: adult male photo
Lesser Spotted Woodpecker: races ID

Lesser Spotted Woodpecker: juvenile ID

Lesser Spotted Woodpecker: adult female ID

Lesser Spotted Woodpecker: adult male ID
Lesser Spotted Woodpecker: general ID

Tuesday, 18 September 2007
Quiz Woodpecker 1

Friday, 14 September 2007
White-backed Woodpecker: juvenile lilfordi race ID

White-backed Woodpecker: female lilfordi race ID
White-backed Woodpecker: male lilfordi race ID
White-backed Woodpecker: lilfordi race general ID
The general features described elsewhere in this blog for nominate leucotos apply equally to the southern European race lilfordi. However, there are some striking and important differences in plumage that together with wing and bill measurements, voice, habitat selection and distribution, have resulted in this race sometimes being considered a separate species i.e. Lilford’s Woodpecker. Lilfordi is slightly larger, overall darker and more heavily streaked with long, black, flecks on the flanks, breast and belly. In addition to vertical streaks there are also some horizontal bars on the flanks that nominate lacks. The pink ventral area extends further up onto the belly than leucotos, often onto the lower breast. Under-parts various shades of white, cream, often dusky or tinged buff and always more marked and darker than leucotos. The forehead, lores, cheeks and throat are also more yellow. The black post-auricular stripe and malar stripe are broader and, where they join to form a T-junction, below the ear-coverts, a larger black area is formed. Face patterns may vary slightly from bird to bird but on all the post-auricular stripe runs over the ear-coverts and finishes much closer to, or even touches, the nape. On leucotos it never touches the nape and a wider white gap is clear. Wings have narrower white bars than on nominate. But above all lilfordi lacks the clear white lower-back and upper rump patch of leucotos race. Lilfordi is thus less "white-backed" than leucotos. It is rather "barred-backed" or perhaps "ladder-backed" due to the white-back and upper-rump being vermiculated with black. The rump of this race often seems totally black.
White-backed Woodpecker: juvenile female ID
Juvenile female White-backeds have red colouration only on the fore-crown (adult female has none) but this often rather patchy, and always less than on juvenile male. Sometimes this patch is merely a few red or pink feathers. The true extent of red on the crown becomes evident some twelve days after hatching.
White-backed Woodpecker: juvenile male ID
Juvenile male White-backeds have an all red crown which extends at the back of the head to the black nape and sideways to the white cheeks. This becomes apparent around twelve days after hatching and is more pronounced as birds peep out of the hole prior to fledging. Male fledglings are generally heavier, and have longer bills and tarsi than sibling females though these factors only noticeable in the hand.
White-backed Woodpecker: juvenile ID

White-backed Woodpecker: adult female ID

White-backed Woodpecker: adult male ID

White-backed Woodpecker: general ID

Middle Spotted Woodpecker: races ID
The nominate race, medius, occurs in Europe though birds on the Greek island of Lesvos may belong to the anatoliae race which breeds on the nearby Turkish mainland. Anatoliae is slightly smaller and more heavily marked on the flanks and breast than medius. But generally differences in size and colour are clinal. As a rule populations in northern and central Europe are larger and paler than in the Balkans and Mediterranean zone.
Middle Spotted Woodpecker: juvenile ID
Black areas of plumage on juvenile Middle Spotted Woodpeckers are less glossy than on adults. Recently fledged birds have brownish greater wing coverts whereas adults have totally black greater wing coverts. White areas of wing-coverts and scapulars have brownish marks. White upper-parts are buffer with some brownish feather tips. White under-parts are greyish rather than yellowish. White on face smudged with grey. Red crown is duller and mottled with grey feather tips. Some red feather tips on hind-crown. Under-tail is a paler, washed out pink and the coloured area is smaller than on adults, not reaching the belly. Some fine, thin barring on flanks, not as heavy markings as adults. Eye grey-brown (in adult dark red). When crown raised, feathers are visibly shorter than on adult. Vent and under-tail coverts washed out, pale pink. Juvenile male have smaller and duller red crowns than adults and the red is flecked with grey or black feathers, especially at the rear and sides. Juvenile females look like juvenile males except the pale red crown patch is often shorter and smaller and the hind-crown is grey-black not golden as adult female. However, many juveniles will be difficult to sex in the field.
Middle Spotted Woodpecker: adult female ID

Middle Spotted Woodpecker: adult male ID

Middle Spotted Woodpecker: general ID

Thursday, 13 September 2007
Woodpecker signs: ringing
Ringing is a series of small, more or less horizontal, holes or grooves which woodpeckers peck around a trunk in order to get at and drink sap. Ringing is sometimes referred to as “girdling" and the peck marks called wells. Ringing is less common amongst European woodpeckers than it is, for example, amongst North American species: there are no true sapsuckers in Europe. Great Spotted, Middle Spotted and Three-toed Woodpeckers are known to ring trees for sap. Syrian, Lesser Spotted and Green Woodpeckers probably do not ring trees themselves though they do take sap from wells made by other. Black Woodpeckers do not ring trees but drink sap from large holes knocked into pines. Woodpeckers ring trees so that sap flows out from the created wells and is mostly done in spring when the sap is rising. The wells pierce the bark and cambial strata of the trunk to tap into the sap conducting routes of the tree. Sunny south-facing trunks, where sap rises first and fastest are often ringed first and most intensely. On productive trees the ringing lines will continue and encircled (girdle) the trunk. Such line may be broken or zigzagged. Several tree species are ringed though pines Pinus and limes Tilia are favourites. Intense ringing can leave wide areas of exposed bark-less trunk. In a given population of Great Spotted Woodpecker most will ring trees but only certain birds will regularly and intensely ring trees.
Woodpecker signs: holes

Woodpecker signs: ruts in turf
Woodpecker signs: cones

The phot here shows a conifer cones wedged into a tree crevice anvil by Great Spotted Woodpecker in Budapest, Hungary (Szabolcs Kókay)
Woodpecker signs: nuts

Woodpecker signs: anvils

Woodpecker signs: dust baths
Dust bathing is not common amongst woodpeckers, probably because of the essentially arboreal lifestyle that most lead, that is, most are reluctant to drop to the ground for any length of time. Indeed, some woodpeckers will not sit on the ground for any reason. However, as with bathing in water, it may be the case that dusting is practised by most woodpeckers but is rarely observed. Black Woodpeckers are known to dust bathe and the more terrestrial Wryneck and Green Woodpecker also occasionally indulges in this activity. Dust baths are often temporary and located on forest tracks. They can be recognised by wing, body and foot prints, though it is often impossible to assign these to one species.
Woodpecker signs: barking?
Many mammals feed on and mark trees: beavers, voles, mice, squirrels, hares, rabbits, deer, even sheep and goats. Generally the marks they leave are easy to separate from those left by woodpeckers, as gnawing and individual tooth marks are left by mammals rather than peck marks. Barking refers to the marks left by deer and elk when they have eaten tree bark. I include it here because such markings can be confused with woodpecker work. Most barking takes place in spring when trees are growing and thus their sap is rising. This of course is also a period of great woodpecker activity. The mammals tear off bark in long strips (usually from bottom up) and leave large incisor marks. In winter bark is tougher and more firmly gripped onto the trunk, so the marks left by deer then are different, with clear lines or furrows left. By contrast woodpecker leave fine peck marks when stripping off bark and the bark is torn off in short sections and from top to bottom.
Woodpecker signs: wood-chips

Wednesday, 12 September 2007
Woodpecker signs: droppings
Finding and identifying woodpecker droppings is one of the great challenges that lovers of the Picidae face. Finding a woodpecker faeces sac in a forest is probably harder than finding the famous needle in a haystack. Woodpecker droppings quickly rot, whereas needles, made as they are from steel, last for ages. Green Woodpecker droppings are the only ones that are likely to be found with any regularity as this species often feed on lawns and areas with flat-cropped turf and so its droppings can be conspicuous, at least for a while. They are greyish, elongated, cylindrical and composed of invertebrate remains, often chitin, dried-out ant bodies and usually enclosed in a membrane, a sac. Such droppings are usually in piles at foraging sites or nearby, and may resemble soggy cigarette butts. They might be confused with small mammal droppings. Green Woodpecker droppings are most often found in winter, when birds do most of their foraging on anthills.
Woodpecker signs: tracks
In Europe only one woodpecker species, Green Woodpecker, is really terrestrial. Grey-headed Woodpecker is partly so, Wryneck and Black occasionally work on the floor when gathering ants, but all other species rarely forage on the ground itself, though some, such as White-backed and Gresat Spotted, often descend to low levels to work on fallen logs. The only realistic situation in which to find woodpecker tracks (i.e. footprints) is after snowfall. Woodpeckers only hop in mud by accident. The first thing to remember, of course, is that all European woodpeckers except Three-toed (sic) have four toes, two pointing forwards and two back. Nine out of the ten European woodpeckers have a syndactyl formation of the toes (the second and third toes point forwards in parallel and toes one and four backwards). Three-toed Woodpecker has the first toe missing, with just toe four pointing backwards and toes two and three forwards). In the rare event that woodpecker tracks are found on the ground (in mud or snow) the quality of the tracks will depend upon the ground conditions.
Woodpecker signs: feeding holes

Woodpecker signs: general

Syrian Woodpecker: female fledgling photo

Syrian Woodpecker: tail pattern

Syrian Woodpecker: juvenile female (fledgling) ID

Syrian Woodpecker: juvenile female ID

Syrian Woodpecker: juvenile male ID
Juvenile Syrian Woodpeckers of both sexes have a black-bordered red crown but the area of red is larger on males, covering most of the crown with a thinner black border. These features are not, however, easy to see in the field.
Syrian Woodpecker: juvenile ID

Syrian Woodpecker: adult female ID

This photo of a female was taken in the Evros Delta, Greece (Bill Baston). Note the lack of red on the nape. The complete white cheek, with no extensive post-auricular stripe, pink undertail and lack of white in the tail are all clearly visible.
Syrian Woodpecker: adult male ID

Syrian Woodpecker: general ID

Tuesday, 11 September 2007
A note on personal observations
I have watched woodpeckers in most European countries, however most of my woodpecker studies have been in Hungary, a country where nine (9) species of woodpecker can be seen. Grey-headed, Green, Black, Great Spotted, Syrian, Middle Spotted, White-backed Woodpecker and Lesser Spotted Woodpeckers are resident and Wryneck occurs from April to August. Three-toed Woodpecker does not occur in Hungary but is resident in all of the neighbouring countries. I regularly observe seven (7) species near my home in Budapest. If I am at home I can observe Syrian and Great Spotted Woodpeckers on an almost daily basis as both frequent our garden!
Sexual dimorphism in woodpeckers

Great Spotted Woodpecker: races ID
There are strong clinal variations in the plumages and sizes of Great Spotted Woodpecker across its range. This is especially so as regards the colour of under-parts. Northern birds are generally larger with stubbier bills and whiter under-parts and foreheads, whilst southern European birds are smaller with slighter bills and often greyish under-parts. Great Spotteds in the British Isles (sometimes claimed as separate race “anglicus" have a slighter, more slender bill and are overall less stocky than the nominate major and continental pinetorum race. They are also matt black, and sometimes rather brownish, on the upper-parts and white areas maybe more creamy or buff. Adult pinetorum from most of mainland Europe is smaller than major, has white under-parts more grey or buff and a longer, thinner bill. Adult hispanus from Iberia is overall darker than most races. White parts are cream, less white on flight feathers and there is sometimes a faint pink flush or a few red blotches on the chest. The under-tail coverts are pinkish, not as bright red as on major. Adult hartertifound on Sardinia and Corsica is a large race with reduced white spots on the wings, very grey under-parts and with a deep red ventral area. Adult canariensis on Tenerife is also a large, dark race and with rather orange-red under-tail coverts and ventral area. Adult thanneri on Gran Canaria is similar to birds on Tenerife but rather pale brown below.
Great Spotted Woodpecker: juvenile ID

Great Spotted Woodpecker: adult female preening

Great Spotted Woodpecker: adult female ID
Subscribe to:
Posts (Atom)